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a LANGE medical book
Harper’s
Illustrated
Biochemistry
twenty-sixth edition
Robert K. Murray, MD, PhD
Professor (Emeritus) of Biochemistry
University of Toronto
Toronto, Ontario
Daryl K. Granner, MD
Joe C. Davis Professor of Biomedical Science
Director, Vanderbilt Diabetes Center
Professor of Molecular Physiology and Biophysics
and of Medicine
Vanderbilt University
Nashville, Tennessee
Peter A. Mayes, PhD, DSc
Emeritus Professor of Veterinary Biochemistry
Royal Veterinary College
University of London
London
Victor W. Rodwell, PhD
Professor of Biochemistry
Purdue University
West Lafayette, Indiana
Lange Medical Books/McGraw-Hill
Medical Publishing Division
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3/16/04 11:10 AM Page ii
Harper’s Illustrated Biochemistry, Twenty-Sixth Edition
Copyright © 2003 by The McGraw-panies, Inc. All rights reserved. Printed in the United States of America. Except as
permitted under the United States Copyright Act of 1976, no part of this publication may be reproduced or distributed in any
form or by any means, or stored in a data base or retrieval system, without the prior written permission of the publisher.
Previous editions copyright © 2000, 1996, 1993, 1990 by Appleton & Lange; copyright © 1988 by Lange Medical Publications.
234567890DOC/DOC09876543
ISBN 0-07-138901-6
ISSN 1043-9811
Notice
Medicine is an ever-changing science. As new research and clinical experience broaden our knowledge, changes in treat-
ment and drug therapy are required. The authors and the publisher of this work have checked with sources believed to be
reliable in their efforts to provide information that plete and generally in accord with the standards accepted at the
time of publication. However, in view of the possibility of human error or changes in medical s