文档介绍：1 !"#$%&'()*' (a) !"#$!.(b) !"%$!.(c) !"&$!. &'(+)',-./ * !" 2)#$$ 1) % 3) & 2) '() 1,2)** ('()*+,-,).,/0'(230009123+,",4056100084123+,78,/056100084) 0$9:; !"<7=>?***@ABCDEF9:;GHIJKLM0NA$O!P !OQRSTUFJKVMW9: ;XYZ[\]^_`abcde fghi9:;jklFm !"nopaq9A$O!rO0sjtuvwxTU F !"y$! zi{ !"LME|}7=P~}7=j]^_`aFbcde0 W!s7=,[\F"#$%D E&'() *! !"y$!+,-.!s/ !"$O!F",0;01DE2Y>3",-4560sE78F"# 9:j;<k7=>? !" ***@AlB7=",PCD7=,&'F-EFG0HIJWKq !"y$!F9:;YL _MFNO3CPPQRST0+UVWXYZ[\]k^ +_` !"y$!F",sdA2Y>Fp ae"0;P",fgh00ijk_O!Yk^FMl",7=,"#bmvnW! !"y$!7=[\F"#F^ 123 !"y$!09:;GHIJK0 !"09:;op 45678 DOI :. 9:(4;9::<./ ProgressinBiochemistryandBiophysics 2014,41(5):413~424 *qrsit,Ouvwxf(21102083). **yz{4H. Tel:**********,E-mail:******@. |}~%2013-06-060U ~%2013-08-19 9:; !"F!""#]G!20#$% &'Levene6 [1](E1906)*kl+,-9:j DE !OQ0 .^1932)/I [2]/051{ !OQ< !"F#$! ******@42 !" $O!3^450tu9Ahi$O!(#$!3 %$!3&$!36$!37$!3h8$!39 $!3:;$!Zy$!)P<=$! !"p atK4AVM0>O- !"3N- !"3S- !"ZwO !"sj !"#$!3 !"% $!P!"&$!?***@BC"#(A1) BIE CDBEPbFj]^_`aFde [3] <0s /***@G!"$O!HI`KJKL[\M-m NO0 !"y$!*<***@P`aBQ !"y$!<***@ABCDEF !"$O !0QRSsE~}CD?KJzi-n !"# $!0T< !"&$!FUVW [4] XYNZP RS[Fh20\] !"y$!q^A-.! OZPRS[TUF!"#$!3 !"%$! K!"&$!F",0; ._09:;jy$ !rOF !"W<***@D?9:;[\F `a9M [5] +_`` !"y$!FklPk^ ab1k0cXs!"7=",0;F"#0iF ***@Gdk{JKI$O!7e[\F!""# 1 =>?@ !"y$!e(*Severin6 [6]!1947)f l",'g1963)0Boyerhy [7]/Eij>9 :;F"#jkgs7"l4 HIe%mn^ !"y$!m\<***@P!"pFjo>pq0X HN COOH PO OO O HN COOH PO OO O HN COOH PO OO O (a) (b) (c) 9:(4;9::<./ . 2014;41(5) r3stF<0W!!"y$!dA2Y> ******@.DECv$wvx0+yCW! !"y$!z{F_|j`2Y>3 }~3- !y$! [20] iX0O!W! !F78"#8 u:0Wq_| [21]`2Y>3 }~1- !y$ !$Y%&0Delbaere6 [22]E9:;'>,F" #z{j(`2 Z3 dP2Y>C)}dN! 1n !"ZN" 2n !"Fy$! ~_*+,-" ,{'2(IUPAC)vnq/7"{'2(IUB){'k Q_"!Kq !OQF3^7=CDF458u :0sj6tK_******@W2Y> [23] E{8u: j02Y>2 @}~3(1)- !y$!0X2Y> 3 @}~1(3)- !y$!* 9phosphor:;~9phosphono:vQ<PDpq Y&0{8u:(45`2Y>2 }~# - !y $!Kpros - !y$!0X2Y>3 }~$ - ! y$!Ktele - !y$! 3